The Biodiversity Ecosystem Functioning experiment in China has been established in 2009. It involves a huge pool of subtropic tree and shrub species. The main goal is determining the role of tree and shrub species diversity for production, element cycling and species conservation in Chinese subtropical forest ecosystems. Novel ecosystem functioning variables highly relevant for environmental issues in China are also investigated, e.g., prevention of soil erosion, carbon sequestration, and resistance to invasion.

One of the BEF-China sites after weeding in 2009 (Photo: Xuefei Yang). Some plots in site A in 2019 (Photo: Xiaojuan Liu).

The experiment consists of two sites (site A, site B), 5 km apart, with two separate species pools that overlap by only 8 species out of the overall tree species pool of 40. All species are equally represented at each diversity level. At one of the two sites, trees of known maternal seed families were planted for 13 tree species.
The plots were planted with a decreasing number of species. The most species-rich plots contain 24 species; plots with 16, 8, 4, 2, 1 and even without trees species were established according to different extinction schemes. In addition, plots with economically important species, plots for free succession and plots with shrub monocultures were planted. Within every plot, 400 trees were planted in a regular fashion (0.6 trees m-2), but mixture of different species was random. In the outer part of the plots, subplots are manipulated to study, e.g., vascular plants in the herb layer, mycorrhiza, decomposers, and herbivores.

Arrangement of the plots at the two sites. The yellow plots are monocultures but also plots planted with commercially important tree species: Cunninghamia lanceolata and Pinus massoniana.

Site characteristics

location Xingangshan
former land use
soil type
area 38.4 ha
no of plots 566
plot size 25.82 m x 25.82 m
no of trees planted 307 200
planting date 2009, 2010
diversity variables species richness
shrub species richness
genetic diversity
diversity gradient 0, 1, 2, 4, 8, 16, 24 tree sp.
2, 4, 8 shrub sp.
size species pool 40 broadleaved tree species + 18 shrub species + 2 commercial coniferous species
species pool tree species: Acer davidii, Ailanthus altissima, Alniphyllum fortune, Betula luminifera, Castanea henryi, Castanopsis carlesii, C. eyrie, C. fargesii, C. sclerophylla, Celtis biondii, Choerospondias axillaris, Cinnamomum camphora, Cyclobalanopsis glauca, C. myrsinifolia, Daphniphyllum oldhamii, Diospyros japonica, Elaeocarpus chinensis, E. glabripetalus, E. japonicas, Idesia polycarpa, Koelreuteria bipinnata, Liquidambar formosana, Lithocarpus glaber, Machilus grijsii, M. leptophylla, M. thunbergii, Manglietia fordiana, Meliosma flexuosa, Nyssa sinensis, Phoebe bournei, Quercus acutissima, Q. fabri, Q. phillyreoides, Q. serrate, Rhus chinensis, Sapindus saponaria, Schima superba, Triadica cochinchinensis, T. sebifera

shrub species: Ardisia crenata, Camellia chekiangoleosa, Distylium buxifolium, D. myricoides, Euonymus myrianthus, Eurya muricata, Ficus erecta, Gardenia jasminoides, Hydrangea chinensis, Itea omeiensis, Loropetalum chinense, Photinia hirsute, Phyllanthus glaucus, Rhaphiolepis indica, Rhododendron ovatum, R. simsii, Syzygium buxifolium, Viburnum setigerum

commercial species: Cunninghamia lanceolata, Pinus massoniana
contact person Xiaojuan Liu


We study the effects of species richness and composition on a range of ecosystem functions, including primary productivity, carbon and nutrient cycling, soil processes, the abundance and biodiversity of other trophic groups, and soil erosion. We also investigate whether genetic diversity affects tree performance, multitrophic interactions, and the response to different species diversity levels or environmental variables such as light availability, which may itself be affected by species diversity.

Extra information

Send an e-mail to the contact person, visit the experiment's own website, or explore the publications: theses

  • Goebes P (2015) Mechanisms of soil erosion in subtropical forests of China - effects of biodiversity, species identity, tree architecture and spatial variability on erosivity. PhD thesis, University of Tübingen
  • Kröber (2015) Stomatal control, xylem hydraulics and leaf morphology in the 40 BEF-China tree species: trait interrelationships, functional diversity and tree growth prediction. PhD thesis, Martin Luther University
  • Cesarz S, Craven D, Auge H, Bruelheide H, Castagneyrol B, Gutknecht J, Hector A, Jactel H, Koricheva J, Messier C, Muys B, Brien MJO, Paquette A, Ponette Q, Potvin C, Reich PB, Scherer-Lorenzen M, Smith AR, Verheyen K, Eisenhauer N (2022) Tree diversity effects on soil microbial biomass and respiration are context dependent across forest diversity experiments. Global Ecology and Biogeography 31:872-885 -
  • Gan H, Li X, Wang Y, Lü P, Ji N, Yao H, Li S, Guo L (2022) Plants Play Stronger Effects on Soil Fungal than Bacterial Communities and Co-Occurrence Network Structures in a Subtropical Tree Diversity Experiment. Microbiology Spectrum -
  • Huang YY, Schuldt A, Hoenig L, Yang B, Liu XJ, Bruelheide H, Ma KP, Schmid B, Niklaus PA (2022) Effects of enemy exclusion on biodiversity-productivity relationships in a subtropical forest experiment. Journal of ecology –
  • Perles-Garcia MD, Kunz M, Fichtner A, Meyer N, Hardtle W, von Oheimb G (2022) Neighbourhood Species Richness Reduces Crown Asymmetry of Subtropical Trees in Sloping Terrain. Remote Sensing 14: 6 –
  • Schnabel F, Liu X, Kunz M, Barry KE, Bongers FJ, Bruelheide H, Fichtner A, Härdtle W, Li S, Pfaff C-T, Schmid B, Schwarz JA, Tang Z, Yang B, Bauhus J, von Oheimb G, Ma K, Wirth C (2021) Species richness stabilizes productivity via asynchrony and drought-tolerance diversity in a large-scale tree biodiversity experiment. Science Advances 7 -
  • Davrinch A, Haider (2021) Intra-specific leaf trait responses to species richness at two different local scales. Basic and Applied Ecology 55: 20-32 -
  • Kwon T, Shibata H, Kepfer-Rojas S, Schmidt IK, Larsen KS, Beier C, Berg B, Verheyen K, Lamarque J-F, Hagedorn F, Eisenhauer N, Djukic I, TeaCompositionNetwork (2021) Effects of climate and atmospheric nitrogen deposition on early to mid-term stage litter decomposition across biomes. Frontiers in Forests and Global Change 4: 678480 -
  • Yang B, Liang Y, Schmid B, Baruffol M, Li Y, He L, Salmon Y, Tian Q, Niklaus PA, Ma K (2021) Soil Fungi Promote Biodiversity–Productivity Relationships in Experimental Communities of Young Trees. Ecosystems -
  • Singavarapu B, Beugnon R, Bruelheide H, Cesarz S, Du J, Eisenhauer N, Guo L, Nawaz A, Wang Y, Xue K, Wubet T (2021) Tree mycorrhizal type and tree diversity shape the forest soil microbiota. Environmental Microbiology -
  • Perles‐Garcia MD, Kunz M, Fichtner A, Härdtle W, von Oheimb G (2021) Tree species richness promotes an early increase of stand structural complexity in young subtropical plantations. Journal of Applied Ecology -
  • Rutten G, Hönig L, Schwaß R, Braun U, Saadani M, Schuldt A, Michalski SG, Bruelheide H, Mommer L (2021) More diverse tree communities promote foliar fungal pathogen diversity, but decrease infestation rates per tree species, in a subtropical biodiversity experiment. Journal of Ecology 109(5): 2068-2080 -
  • Saadani M, Hönig L, Bien S, Koehler M, Rutten G, Wubet T, Braun U, Bruelheide H (2021) Local tree diversity suppresses foliar fungal infestation and decreases morphological but not molecular richness in a young subtropical forest. Journal of Fungi 7: 173 -
  • Germany MS, Bruelheide H, Erfmeier A, Wagg C (2021) Drivers of understorey biomass: tree species identity is more important than richness in a young forest. Journal of Plant Ecology 14(3): 465-477 -
  • Wang M-Q, Yan C, Luo A, Li Y, Chesters D, Qiao H-J, Chen J-T, Zhou Q-S, Ma K, Bruelheide H, Schuldt A, Zhang Z, Zhu C-D (2021) Phylogenetic relatedness, functional traits, and spatial scale determine herbivore co-occurrence in a subtropical forest. Ecological Monographs 92(1): e01492 -
  • Weinhold A, Döll S, Liu M, Schedl A, Pöschl Y, Xu X, Neumann S, van Dam NM (2021) Tree species richness differentially affects the chemical composition of leaves, roots and root exudates in four subtropical tree species. Journal of Ecology 110(1): 97-116 -
  • Wu D, Staab M, Yu M (2021) Canopy closure retards fine wood decomposition in subtropical regenerating forests. Ecosystems
  • Wu D, Pietsch KA, Staab M, Yu M, (2021) Wood species identity alters dominant factors driving fine wood decomposition along a tree diversity gradient in subtropical plantation forests. Biotropica 53(2): 643-657 -
  • Gheyret G, Zhang H, Guo Y, Liu T, Bai Y, Li S, Schmid B, Bruelheide H, Ma K, Tang Z (2021) Radial growth response of trees to seasonal soil humidity in a subtropical forest. Basic and Applied Ecology -
  • Guo P, Wang M, Orr M, Li Y, Chen J, Zhou Q, Staab M, Fornoff F, Chen G, Zhang N, Klein A-M, Zhu C (2021) Tree diversity promotes predatory wasps and parasitoids but not pollinator bees in a subtropical experimental forest. Basic and Applied Ecology 53:134-142 -
  • Trogisch S, Liu X, Rutten G, Xue K, Bauhus J, Brose U, Bu W, Cesarz S, Chesters D, Connolly J, Cui X, Eisenhauer N, Guo L, Haider S, Härdtle W, Kunz M, Liu L, Ma Z, Neumann S, Sang W, Schuldt A, Tang Z, van Dam NM, von Oheimb G, Wang M-Q, Wang S, Weinhold A, Wirth C, Wubet T, Xu X, Yang B, Zhang N, Zhu C-D, Ma K, Wang Y, Bruelheide H (2021) The significance of tree-tree interactions for forest ecosystem functioning. Basic and Applied Ecology -
  • Li X, Qian X, Gao C, Seitz S, Scholten T, Wang Y, Yao H, Gan H, Guo L (2021) Plant identity strongly structures the root-associated fungal community in a diverse subtropical forest. Basic and Applied Ecology -
  • Hildebrand M, Perles-Garcia MD, Kunz M, Härdtle W, von Oheimb G, Fichtner A (2021) Tree-tree interactions and crown complementarity: The role of functional diversity and branch traits for canopy packing. Basic and Applied Ecology 50: 217-227 -
  • Ma L, Bongers FJ, Li S, Tang T, Yang B, Ma K, Liu X (2021) Species identity and composition effects on community productivity in a subtropical forest. Basic and Applied Ecology.
  • Jansen K, von Oheimb G, Bruelheide H, Härdtle W, Fichtner A (2021) Tree species richness modulates water supply in the local tree neighbourhood: evidence from wood δ13C signatures in a large-scale forest experiment. Proceedings of the Royal Society B-Biological Sciences 288: 20203100 -
  • Bongers FJ (2020) Functional-structural plant models to boost understanding of complementarity in light capture and use in mixed-species forests. Basic and Applied Ecology 48: 92-101 -
  • Zhang N, Bruelheide H, Li Y, Liang Y, Wubet T, Trogisch S, Ma K (2020) Community and neighbourhood tree species richness effects on fungal species in leaf litter. Fungal Ecology 47 -
  • Trogisch S, Albert G, Du J, Wang Y, Xue K, Bruelheide H (2020) Promoting resilience of large international collaborative research programs in times of global crisis. Academic Practice in Ecology and Evolution 10: 12549-12554 -
  • Wang M-Q, Li Y, Chesters D, Bruelheide H, Ma K, Guo P-F, Zhou Q-S, Staab M, Zhu C-D, Schuldt A (2020) Host functional and phylogenetic composition rather than host diversity structure plant‐herbivore networks. Molecular Ecology 29: 2747–2762 -
  • Bongers FJ, Schmid B, Durka W, Li S, Bruelheide H, Hahn CZ, Yan H, Ma K, Liu X (2020) Genetic richness affects trait variation but not community productivity in a tree diversity experiment. New Phytologist 227:744-756 -
  • Bongers FJ, Schmid B, Sun Z, Li Y, Härdtle W, von Oheimb G, Li Y, Li S, Staab M, Ma K, Liu X(2020) Growth-trait relationships in subtropical forest are stronger at higher diversity. Journal of Ecology 108:256-266 -
  • Chen Y, Huang Y, Niklaus PA, Castro-Izaguirre N, Clark AT, Bruelheide H, Ma K, Schmid B (2020) Directed species loss reduces community productivity in a subtropical forest biodiversity experiment. Nature Ecology & Evolution 4:550–559 -
  • Fichtner A, Schnabel F, Bruelheide H, Kunz M, Mausolf K, Schuldt A, Härdtle W, von Oheimb G (2020) Neighbourhood diversity mitigates drought impacts on tree growth. Journal of Ecology 108:865-875 -
  • Skarbek CJ, Noack M, Bruelheide H, Härdtle W, von Oheimb G, Scholten T, Seitz S, Staab M (2020) A tale of scale: Plot but not neighbourhood tree diversity increases leaf litter ant diversity. Journal of Animal Ecology 89: 299–308 -
  • Bruelheide H, Chen Y, Huang Y, Ma K, Niklaus PA, Schmid B (2019) Response to Comment on "Impacts of species richness on productivity in a large-scale subtropical forest experiment". Science 363:eaav9863 -
  • Du Y, Yang B, Chen S, Ma K (2019) Diverging shifts in spring phenology in response to biodiversity loss in a subtropical forest. Journal of Vegetation Science 30:1175-1183 -
  • Fornoff F, Klein A-M, Bluthgen N, Staab M (2019) Tree diversity increases robustness of multi-trophic interactions. Proceedings of the Royal Society B-Biological Sciences. 286:20182399 -
  • Forrester DI, Rodenfels P, Haase J, Härdtle W, Leppert KN, Niklaus PA, von Oheimb G, Scherer-Lorenzen M, Bauhus J (2019) Tree-species interactions increase light absorption and growth in Chinese subtropical mixed-species plantations. Oecologia -
  • Germany MS, Bruelheide H, Erfmeier A (2019) Janzen-Connell effects in a forest BEF experiment: Strong distance-dependent seedling establishment of multiple species. Ecology 100:e02736 -
  • Kunz M, Fichtner A, Härdtle W, Raumonen P, Bruelheide H, von Oheimb G (2019) Neighbour species richness and local structural variability modulate aboveground allocation patterns and crown morphology of individual trees. Ecology Letters 22: 2130-2140 -
  • Li Y, Bruelheide H, Scholten T, Schmid B, Sun Z, Zhang N, Bu W, Liu X, Ma K (2019) Early positive effects of tree species richness on soil organic carbon accumulation in a large-scale forest biodiversity experiment. Journal of Plant Ecology 12:882-893 -
  • Rentschler T, Gries P, Behrens T, Bruelheide H, Kühn P, Seitz S, Shi X, Trogisch S, Scholten T, Schmidt K (2019) Comparison of catchment scale 3D and 2.5D modelling of soil organic carbon stocks in Jiangxi Province, PR China. PloS one 14:e0220881 -
  • Ristok C, Leppert KN, Scherer-Lorenzen M, Niklaus PA, Bruelheide H (2019) Soil macrofauna and leaf functional traits drive the decomposition of secondary metabolites in leaf litter. Soil Biology & Biochemistry 135:429-437 -
  • Schuldt A, Ebeling A, Kunz M, Staab M, Guimaraes-Steinicke C, Bachmann D, Buchmann N, Durka W, Fichtner A, Fornoff F, Härdtle W, Hertzog LR, Klein A-M, Roscher C, Schaller J, von Oheimb G, Weigelt A, Weisser W, Wirth C, Zhang J, Bruelheide H, Eisenhauer N (2019) Multiple plant diversity components drive consumer communities across ecosystems. Nature Communications 10:1460 -
  • Song Z, Seitz S, Li J, Goebes P, Schmidt K, Kühn P, Shi X, Scholten T (2019) Tree diversity reduced soil erosion by affecting tree canopy and biological soil crust development in a subtropical forest experiment. Forest Ecology and Management 444:69-77 -
  • Wang M, Li Y, Chesters D, Anttonen P, Bruelheide H, Chen J, Durka W, Guo P, Härdtle W, Ma K, Michalski SG, Schmid B, von Oheimb G, Wu C, Zhang N, Zhou Q, Schuldt A, Zhu C (2019) Multiple components of plant diversity loss determine herbivore phylogenetic diversity in a subtropical forest experiment. Journal of Ecology 107:2697-2712 -
  • Weißbecker C, Heintz-Buschart A, Bruelheide H, Buscot F, Wubet T (2019) Linking Soil Fungal Generality to Tree Richness in Young Subtropical Chinese Forests. Microorganisms 7:547 -
  • Cao HX, Klein AM, Zhu CD, Staab M, Durka W, Fischer M, Fornoff F (2018) Intra- and interspecific tree diversity promotes multitrophic plant-Hemiptera-ant interactions in a forest diversity experiment. Basic and Applied Ecology 29: 89-97–
  • Djukic I, Kepfer-Rojas S, Kappel Schmidt I, Steenberg Larsen K, Beier C, Berg B, Verheyen K, TeaComposition (2018) Early stage litter decomposition across biomes. Science of the Total Environment
  • Fichtner A, Härdtle W, Bruelheide H, Kunz M, Li Y, von Oheimb G (2018) Neighbourhood interactions drive overyielding in mixed-species tree communities. Nature communications 9: 1144 -
  • Grossman JJ, Vanhellemont M, Barsoum N, Bauhus J, Bruelheide H, Castagneyrol B, Cavender-Bares J, Eisenhauer N, Ferlian O, Gravel D, Hector A, Jactel H, Kreft H, Mereu S, Messier C, Muys B, Nock C, Paquette A, Parker J, Perring MP, Ponette Q, Reich PB, Schuldt A, Staab M, Weih M, Zemp DC, Scherer-Lorenzen M, Verheyen K (2018) Synthesis and future research directions linking tree diversity to growth, survival, and damage in a global network of tree diversity experiments. Environmental and Experimental Botany - doi: 10.1016/j.envexpbot.2017.12.015
  • Huang Y, Chen Y, Castro-Izaguírre, Baruffol M, Brezzi M, Lang A, Li Y, Härdtle W, von Oheimb G, Yang X, Liu X, Pei K, Both S, Yang B, Eichenberg D, Assmann T, Bauhus J, Behrens T, Buscot F, Chen X-Y, Chesters D, Ding B-Y, Durka W, Erfmeiere A, Fang J, Fischer M, Guo L-D, Guo D, Gutknecht JLM, He J-S, He C-L, Hector A, Hönig L, Hu R-Y, Klein A-M, Kühn P, Liang Y, Li S, Michalski S, Scherer-Lorenzen M, Schmidt K, Scholten T, Schuldt A, Shi X, Tan M-Z, Tang Z, Trogisch S, Wang Z, Welk E, Wirth C, Wubet T, Xiang W, Yu M, Yu X-D, Zhang J, Zhang S, Zhang N, Zhou H-Z, Zhu C-D, Zhu L, Bruelheide H, Ma K, Niklaus PA, Schmid B (2018) Impacts of species richness on productivity in a large-scale subtropical forest experiment. Science 362: 80-83 - doi: 10.1126/science.aat6405
  • Huang Y, Ma K, Niklaus PA, Schmid B (2018) Leaf-litter overyielding in a forest biodiversity experiment in subtropical China. Forest Ecosystems 5(38) -
  • Koricheva J, Hayes D (2018) The relative importance of plant intraspecific diversity in structuring arthropod communities: A meta-analysis. Functional Ecology 32: 1704-1717 - doi: 10.1111/1365-2435.13062
  • Song ZS, Seitz S, Zhu PP, Goebes P, Shi XZ, Xu SX, Wang MY, Schmidt K, Scholten T (2018) Spatial distribution of LAI and its relationship with throughfall kinetic energy of common tree species in a Chinese subtropical forest plantation. Forest Ecology and Management 425: 189-195 –
  • Weißbecker C, Wubet T, Lentendu G, Kühn P, Scholten T, Bruelheide H, Buscot F (2018) Experimental evidence of functional group-dependent effect of tree diversity on soil fungi in subtropical forests. Frontiers in Microbiology 9: 2312 - doi: 10.3389/fmicb.2018.02312
  • Bu W, Liu X, Schmid B, Li Y, Härdtle W, von Oheimb G, Liang Yu, Sun Z, Huang Y, Bruelheide H, Ma K (2017) Interspecific and intraspecific variation in specific root length drives aboveground biodiversity effects in young experimental forest stands. Journal of Plant Ecology 10: 158-169 - doi: 10.1093/jpe/rtw096
  • Eichenberg D, Pietsch K, Meister C, Ding W, Yu M, Wirth C (2017) The effect of microclimate on wood decay is indirectly altered by tree species diversity in a litterbag study. Journal of Plant Ecology 10: 170-178 - doi: 10.1093/jpe/rtw116
  • Fichtner A, Härdtle W, Li Y, Bruelheide H, Kunz, M, von Oheimb G (2017) From competition to facilitation: how tree species respond to neighbourhood diversity. Ecology Letters 20: 892-900 - doi: 10.1111/ele.12786
  • Germany M, Bruelheide H, Erfmeier A (2017) Limited tree diversity effects on herb layer composition, richness and productivity in experimental forest stands. Journal of Plant Ecology 10: 190-200 - doi: 10.1093/jpe/rtw109
  • Hahn CZ, Niklaus PA, Bruelheide H, Michalski SG, Shi M, Yang X, Zeng X, Fischer M, Durka W (2017) Opposing intraspecific vs. interspecific diversity effects on herbivory and growth in subtropical experimental tree assemblages. Journal of Plant Ecology 10: 242-251 - doi: 10.1093/jpe/rtw098
  • Leppert KN, Niklaus PA, Scherer-Lorenzen M (2017) Does species richness of subtropical tree leaf litter affect decomposition, nutrient release, transfer and subsequent uptake by plants? Soil Biology and Biochemistry 115: 44-53 - doi: 10.1016/j.soilbio.2017.08.007
  • Li Y, Kröber W, Bruelheide H, Härdtle W, von Oheimb G (2017) Crown and leaf traits as predictors for species-specific tree sapling growth rates in subtropical plantations. Journal of Plant Ecology 10: 136-145 - doi: 10.1093/jpe/rtw041
  • Niklaus PA, Baruffol M, He J-S, Ma K, Schmid B (2017) Can niche plasticity promote biodiversity-productivity relationships through increased complementarity? Ecology 98: 1104-1116 - doi: 10.1002/ecy.1748
  • Peng S, Schmid B, Haase J, Niklaus PA (2017) Leaf area increases with species richness in young experimental stands of subtropical trees. Journal of Plant Ecology 10: 128-135 - doi: 10.1093/jpe/rtw016
  • Ristok C, Leppert K, Franke K, Scherer-Lorenzen M, Niklaus PA, Wessjohann L, Bruelheide H (2017) Leaf litter diversity positively affects the decomposition of plant polyphenols. Plant and Soil 419: 305-317 - doi: 10.1007/s11104-017-3340-8
  • Schmid B, Baruffol M, Wang ZH, Niklaus PA (2017) A guide to analyzing biodiversity experiments. Journal of Plant Ecology 10: 91-110 - doi: 10.1093/jpe/rtw107
  • Scholten T, Goebes P, Kühn P, Seitz S, Assmann T, Bauhus J, Bruelheide H, Buscot F, Erfmeier A, Fischer M, Härdtle W, He J-S, Ma K, Niklaus PA, Scherer-Lorenzen M, Schmid B, Shi X, Song Z, von Oheimb G, Wirth C, Wubet T, Schmidt K (2017) On the combined effect of soil fertility and topography on tree growth in subtropical forest ecosystems - a study from SE China Journal of Plant Ecology 10: 111-127 - doi: 10.1093/jpe/rtw065
  • Schuldt A, Fomoff F, Bruelheide H, Klein A-M, Staab M (2017) Tree species richness attenuates the positive relationship between mutualistic ant-hemipteran interactions and leaf chewer herbivory. Proceedings Royal Society B 284: 20171489 - doi: 10.1098/rspb.2017.1489
  • Schuldt A, Hönig L, Li Y, Fichtner A, Härdtle, von Oheimb G, Welk E, Bruelheide H (2017) Herbivore and pathogen effects on tree growth are additive, but mediated by tree diversity and plant traits. Ecology and Evolution 7: 7462-7474 - doi: 10.1002/ece3.3292
  • Seitz S, Nebel M, Goebes P, Käppeler K, Schmidt K, Shi X, Song Z, Webber CL, Weber B, Scholten T (2017) Bryophyte-dominated biological soil crusts mitigate soil erosion in an early successional Chinese subtropical forest. Biogeosciences 14: 5775-5788 -
  • Staab M, Fornoff F, Klein A-M, Blüthgen N (2017) Ants at Plant Wounds: A Little-Known Trophic Interaction with Evolutionary Implications for Ant-Plant Interactions. American Naturalist 190(3): 442-450 –
  • Staab M, Methorst J, Peters J, Blüthgen N, Klein AM (2017) Tree diversity and nectar composition affect arthropod visitors on extrafloral nectaries in a diversity experiment. Journal of Plant Ecology 10: 201-212 - doi: 10.1093/jpe/rtw017
  • Sun Z, Liu X, Schmid B, Bruelheide H, Bu W, Ma K (2017) Positive effects of tree species richness on fine-root production and biomass. Journal of Plant Ecology 10: 146-157 - doi: 10.1093/jpe/rtw094
  • Trogisch S, Schuldt A, Bauhus J, Blum JA, Both S, Buscot F, Castro-Izaguirre N, Chesters D, Durka W, Eichenberg D, Erfmeier A, Fischer M, Geißler C, Germany MS, Goebes P, Gutknecht J, Hahn CZ, Haider S, Härdtle W, He J-S, Hector A, Hönig L, Huang Y, Klein A-M, Kühn P, Kunz M, Leppert KN, Li Y, Liu X, Niklaus PA, Pei Z, Pietsch KA, Prinz R, Proß T, Scherer-Lorenzen M, Schmidt K, Scholten T, Seitz S, Song Z, Staab M, von Oheimb G, Weißbecker C, Welk E, Wirth C, Wubet T, Yang B, Yang X, Zhu C-D, Schmid B, Ma K, Bruelheide H (2017) Toward a methodical framework for comprehensively assessing forest multifunctionality. Ecology and Evolution 7: 10652-10674 - doi: 10.1002/ece3.3488
  • Wang C, Ma Y, Trogisch S, Huang Y, Geng Y, Scherer-Lorenzen M, He J-S (2017) Soil respiration is driven by fine root biomass along a forest chronosequence in subtropical China. Journal of Plant Ecology 10: 36-46 - doi: 10.1093/jpe/rtw044
  • Yang B, Ying L, Ding B, Both S, Erfmeier A, Härdtle W, Ma K, Schmid B, Scholten T, Seidler G, von Oheimb G, Yang X, Bruelheide H (2017) Impact of tree diversity and environmental conditions on the survival of shrub species in a forest biodiversity experiment in subtropical China. Journal of Plant Ecology 10: 179-189 - doi: 10.1093/jpe/rtw099
  • Yang B, Li B, He Y, Zhang L, Bruelheide H, Schuldt A (2017) Tree diversity has contrasting effects on predation rates by birds and arthropods on three broadleaved, subtropical tree species. Ecological Research 33(1): 205-2012 -
  • Zhang J, Bruelheide H, Chen X, Eichenberg D, Kröber W, Xu X, Xu L, Schuldt A (2017) Tree diversity promotes generalist herbivore community patterns in a young subtropical forest experiment. Oecologia 183: 455-467 - doi: 10.1007/s00442-016-3769-0
  • Goebes P, Schmidt K, Härdtle W, Seitz S, Stumpf F, von Oheimb G, Scholten T (2016) Rule-based analysis of throughfall kinetic energy to evaluate biotic and abiotic factor thresholds to mitigate erosive power. Progress in Physical Geography 40: 431-449 - doi: 10.1177/0309133315624642
  • Pei Z, Eichenberg D, Bruelheide H, Kröber W, Kühn P, Li Y, von Oheimb G, Purschke O, Scholten T, Buscot F, Gutknecht JLM (2016) Soil and tree species traits both shape soil microbial communities during early growth of Chinese subtropical forests. - Soil Biology and Biochemistry 96: 180-190 - doi: 10.1016/j.soilbio.2016.02.004
  • Purahong W, Durka W, Fischer M, Dommert S, Schöps R, Buscot F, Wubet T (2016) Tree species, tree genotypes and tree genotypic diversity levels affect microbe-mediated soil ecosystem functions in a subtropical forest. Scientific Reports 6: 36672 - doi: 10.1038/srep36672
  • Seidelmann KN, Scherer-Lorenzen M, Niklaus PA (2016) Direct vs. microclimate-driven effects of tree species diversity on litter decomposition in young subtropical forest stands. Plos ONE 11(8): e0160569 - doi: 10.1371/journal.pone.0160569
  • Seitz S, Goebes P, Song Z, Bruelheide H, Härdtle W, Kühn P, Li Y, Scholten T (2016) Tree species and functional traits but not species richness affect interrill erosion processes in young subtropical forests. Soil 2: 49-61 - doi: 10.5194/soil-2-49-2016
  • Verheyen K, Vanhellemont M, Auge H, Baeten L, Baraloto C, Barsoum N, Bilodeau-Gauthier S, Bruelheide H, Castagneyrol B, Godbold D, Haase J, Hector A, Jactel H, Koricheva J, Loreau M, Mereu S, Messier C, Muys B, Nolet P, Paquette A, Parker J, Perring M, Ponette Q, Potvin C, Reich P, Smith A, Weih M, Scherer-Lorenzen M (2016) Contributions of a global network of tree diversity experiments to sustainable forest plantations. Ambio 45: 29-41 - doi: 10.1007/s13280-015-0685-1
  • Goebes P, Bruelheide H, Härdtle W, Kröber W, Kühn P, Li Y, Seitz S, von Oheimb G, Scholten T (2015) Species-specific effects on throughfall kinetic energy in subtropical forest plantations are related to leaf traits and tree architecture. PLoS ONE 10: e0128084 - doi:10.1371/journal.pone.0128084
  • Goebes P, Seitz S, Kühn P, Li Y, Niklaus P, von Oheimb G, Scholten T (2015) Throughfall kinetic energy in young subtropical forests: Investigation on tree species richness effects and spatial variability. Agricultural and Forest Meteorology 213: 148-159 - doi: 10.1016/j.agrformet.2015.06.019
  • Kröber W, Heklau H, Bruelheide H (2015) Leaf morphology of 40 evergreen and deciduous broadleaved subtropical tree species and relationships to functional ecophysiological traits. Plant Biology 17: 373-383 - doi: 10.1111/plb.12250
  • Kröber W, Li Y, Härdtle W, Ma K, Schmid B, Schmidt K, Scholten T, Seidler G, von Oheimb G, Welk E, Wirth C, Bruelheide H (2015) Early subtropical forest growth is driven by community mean trait values and functional diversity rather than the abiotic environment Ecology and Evolution 5: 3541-3556 - doi: 10.1002/ece3.1604
  • Kröber W, Plath I, Heklau H, Bruelheide H (2015) Relating stomatal conductance to leaf functional traits. Journal of Visualized Experiments e52738 - doi: 10.3791/52738 -
  • Paine CET, Amissah L, Auge H, Baraloto C, Baruffol M, Bourland N, Bruelheide H, Daïnou K, de Gouvenain RC, Doucet J-L, Doust S, Fine PVA, Fortunel C, Haase J, Holl KD, Jactel H, Li X, Kitajima K, Koricheva J, Martínez-Garza C, Messier C, Paquette A, Philipson C, Piotto D, Poorter L, Posada JM, Potvin C, Rainio K, Russo SE, Ruiz-Jaen M, Scherer-Lorenzen M, Webb CO, Wright SJ, Zahawi RA, Hector A (2015) Globally, functional traits are weak predictors of juvenile tree growth, and we do not know why. Journal of Ecology 103: 978-989 - doi: 10.1111/1365-2745.12401
  • Pfaff C-T, König-Ries B, Lang AC, Ratcliffe S, Wirth C, Man X, Nadrowski K (2015) rBEFdata: documenting data exchange and analysis for a collaborative data management platform. Ecology and Evolution 5: 2890-2897 - doi: 10.1002/ece3.1547
  • Schuldt A, Bruelheide H, Härdtle W, Assmann T, Li Y, Ma K, von Oheimb G, Zhang J (2015) Early positive effects of tree species richness on herbivory in a large-scale forest biodiversity experiment influence tree growth. Journal of Ecology 103: 563-571 - doi: 10.1111/1365-2745.12396
  • Staab M, Blüthgen N, Klein AM (2015) Tree diversity alters the structure of a tri-trophic network in a biodiversity experiment. Oikos 124: 827-834 - doi: 10.1111/oik.01723
  • Bruelheide H, Nadrowski K, Assmann T, Bauhus J, Both S, Buscot F, Chen X-Y, Ding B, Durka W, Erfmeier A, Gutknecht JLM, Guo D, Guo L-D, Härdtle W, He J-S, Klein A-M, Kühn P, Liang Y, Liu X, Michalski S, Niklaus P, Pei K, Scherer-Lorenzen M, Scholten T, Schuldt A, Seidler G, Trogisch S, von Oheimb G, Welk E, Wirth C, Wubet T, Yang X, Yu M, Zhang S, Zhou H, Fischer M, Ma K, Schmid B (2014) Designing forest biodiversity experiments: general considerations illustrated by a new large experiment in subtropical China. Methods in Ecology and Evolution 5: 74-89 - doi: 10.1111/2041-210X.12126
  • Kröber W, Bruelheide H (2014) Transpiration and stomatal control: a cross-species study of leaf traits in 39 evergreen and deciduous broadleaved subtropical tree species. Trees - Structure and Function 28: 901-914 - doi: 10.1007/s00468-014-1004-3
  • Kröber W, Zhang SR, Ehmig M, Bruelheide H (2014) Linking xylem hydraulic conductivity and vulnerability to the leaf economics spectrum - A cross-species study of 39 evergreen and deciduous broadleaved subtropical tree species. PLoS ONE 9: e109211 - doi: 10.1371/journal.pone.0109211
  • Li Y, Hess C, von Wehrden H, Härdtle W, von Oheimb G (2014) Assessing tree dendrometrics in young regenerating plantations using terrestrial laser scanning. Annals of Forest Science 71: 453-462 - doi: 10.1007/s13595-014-0358-4
  • Li Y, Härdtle W, Bruelheide H, Nadrowski K, Scholten T, von Wehrden H, von Oheimb G (2014) Site and neighborhood effects on growth of tree saplings in subtropical plantations (China). Forest Ecology and Management 327: 118-127 - doi: 10.1016/j.foreco.2014.04.039
  • Nadrowski K, Ratcliffe S, Bönisch G, Bruelheide H, Kattge J, Liu X, Maicher L, Mi X, Prilop M, Seifarth D, Welter K, Windish S, Wirth C (2013) Harmonizing, annotating and sharing data in biodiversity-ecosystem functioning research. Methods in Ecology and Evolution 4: 201-205 - doi: 10.1111/2041-210x.12009
  • Yang X, Bauhus J, Both S, Fang T, Härdtle W, Kröber W, Ma K, Nadrowski K, Pei K, Scherer-Lorenzen M, Scholten T, Seidler G, Schmidt B, von Oheimb G, Bruelheide H (2013) Establishment success in a forest biodiversity and ecosystem functioning experiment in subtropical China (BEF-China) European Journal of Forest Research 132: 593-606 - doi: 10.1007/s10342-013-0696-z

photo plot photo plot
Site A - plot M27 in 2014 Site B in 2019